Adaptive
immunity against pathogen challenge is organised within lymph nodes (LNs),
small organs that enable constant screening of tissue-derived material by
circulating lymphocytes in designated compartments and facilitate the
establishment of immune responses. A central feature of their specialised
microanatomy is an intricate network of blood and lymphatic channels, which
serve as entry and exit routes for immune cells and lymph, thereby orchestrating
the course of immune cell migration and antigen transmission. However, we
currently lack understanding of the global organisation of both vascular
systems and their structural dynamism during inflammation, when LNs increase rapidly
in size in response to changes in cell trafficking. To gain better insight into
the spatial organisation of blood and lymphatic systems, we aimed to generate
high-resolution images of entire LNs and describe the vascular topology with
the help of sophisticated computer analysis and visualisation tools. Using a
unique automated confocal imaging system we generated unprecedented 3D images
of blood and lymphatic channels in murine LNs at subcellular resolution, which
provided new insights into their labyrinthine arrangement and regional specialisation.
Moreover, we developed custom image-processing and analysis tools to assess the
vascular volumes and distances between blood and lymphatic channels. Detailed
quantification of the blood vessel network yielded information about vessel
diameters, lengths, and densities in 2D and 3D. In addition, we established
methods to compare vessel parameters in LN subregions such as the T and B cell
zones and evaluated the distribution of HEVs. In a first attempt to quantify
changes in reactive LNs we analysed the vascular systems of an LPS-stimulated
LN and found the blood vasculature significantly expanded after just two days
compared to unstimulated controls. These methods can now be used to further investigate
vascular remodelling processes during LN activation, and will help us to map LN
anatomy more comprehensively.